Plants, which had begun colonizing the land during the Silurian Period, continued to make evolutionary progress during the Devonian. Lycophytes, horsetails and ferns grew to large sizes and formed Earth’s first forests. By the end of the Devonian, progymnosperms such as Archaeopteris were the first successful trees. Archaeopteris could grow up to 98 feet (30 meters) tall with a trunk diameter of more than 3 feet. It had a softwood trunk similar to modern conifers that grew in sequential rings. It did not have true leaves but fern-like structures connected directly to the branches (lacking the stems of true leaves). There is evidence that they were deciduous, as the most common fossils are shed branches. Reproduction was by male and female spores that are accepted as being the precursors to seed-bearing plants. By the end of the Devonian Period, the proliferation of plants increased the oxygen content of the atmosphere considerably, which was important for development of terrestrial animals. At the same time carbon dioxide (CO2), a greenhouse gas, was depleted from earlier levels. This may have contributed to the cooling climate and the extinction event at the end of the Devonian.
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